TYPE: Costa Rica. Turrialba: Tuis, Cien Manzanas, 9°51′25″N 83°33′47″W, ca. 3,117 feet (950 m), along a minor tributary of Río Pacuare, premontane wet forest, secondary vegetation and remnants of primary, May 26, 2003, F. Pupulin 4801, R. and K. Dressler, J. Aguilar, G. Gerlach, P. Kindlmann, H. León-Paéz, S. Pugh-Jones & E. Serrano (holotype, JBL-Spirit; illustration of type).